Pseudoscorpiones

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Pseudoscorpiones de Geer, 1778
Pseudoskorpione
Neobisium sp IMG 0705bepr (Medium) Oertel.JPG
Neobisium sp.
Systematik
Klasse: Arachnida (Spinnentiere)

Pseudoscorpiones ist eine Ordnung der Klasse Arachnida. Weltweit kommen 1100 Arten vor. Die größte Art ist Garypus giganteus, welche 7 mm lang wird. (Kaestner 1956)

Sie werden im deutschen Sprachraum Pseudoskorpione oder auch Afterskorpione genannt. (Kaestner 1956)

Determination

→ siehe Artikel Familienschlüssel Pseudoskorpione

Anatomie

Pseudoscorpiones sind sehr kleine Tiere. Sie haben eine durchschnittliche Körperlänge von 2 bis 4 mm. Die kleinsten Arten sind kleiner als 1 mm. (Kaestner 1956) Die Vertreter der Ordnung ähneln den Skorpionen, besitzen aber weder Schwanz noch Endstachel. Der Körper ist in Prosoma (oder Cephalothorax) und Opisthosoma (Abdomen) geteilt, die breit miteinander verbunden sind. Das Prosoma ist oben vom Carapax bedeckt, der nicht geteilt ist. Die Unterseite des Cephalothorax zeigt jedoch eine deutliche Segmentierung. Das Opisthosoma ist in 11 oder 12 Glieder geteilt. Pseudoskorpione haben scherenähnliche Pedipalpen, die oft auch Giftdrüsen aufweisen. Die Cheliceren besitzen oft Spinndrüsen. Pseudoskorpione haben 4 Laufbeinpaare. (Parker 1982)

→ siehe auch Hauptartikel Anatomie der Pseudoskorpione

Lebensweise

Zur Fortpflanzung legt das Männchen die Spermazellen in ein Spermatophor, und das Weibchen nimmt sie auf. Einige Arten leben als Zwitter. Alle bekannten Arten leben räuberisch. Manche Arten heften sich gelegentlich an Insekten (vor allem die trächtigen Weibchen) und lassen sich so transportieren (Phorese). (Parker 1982)

Lebensraum

Pseudoscorpiones leben vor allem in der Streu, in Bodenlücken, unter Baumrinde, in Kompost- und Heuhaufen, in Höhlen, kommen aber auch synanthrop vor. (Blick & Muster 2003) Einige Arten sind an Meeresküsten heimisch, sind aber nicht wirklich aquatil. (Parker 1982)

Verbreitung

Die Ordnung ist weltweit verbreitet (auf dem antarktischen Festland nicht, aber zwei Arten sind von den französischen subantarktischen Inseln bekannt). (Parker 1982)

Familienspektrum

In Europa und im Maghreb kommen nach Datenlage dieses Wikis 14 Familien der Ordnung Pseudoscorpiones vor.[A]

NameNOSEFIIEGBGB-NIRDKDENLBELUPLLTLVEECHATLICZSKHU
Atemnidae     ?      ?  ××? ××
Cheiridiidae×××××?×××× ×?××××?×××
Cheliferidae×××××?×××× ×?××××?×××
Chernetidae×××××?×××××××××××?×××
Chthoniidae×××××?××××××? ×××?×××
Geogarypidae     ?      ?   ×?   
Larcidae ×  ×?×××  ×?×  ×?×××
Neobisiidae×××××?××××××?××××?×××
Syarinidae×    ? ×    ?  ××?×  
Withiidae    ×?×××   ?  ××? ××
NamePTESES-IBADFRFRHITIT82IT88ROBGBASIHRMEMKRSKVALGRGR-M
Atemnidae×   × ×××××  ×    ×× 
Bochicidae×                    
Cheiridiidae×   × × ×××  ××   ×× 
Cheliferidae××  ×××××××× ×××× ×× 
Chernetidae×   ×××××××××××××××× 
Chthoniidae×××××××××××××××××××× 
Garypidae×   × ×××  × ×     × 
Garypinidae×   × ×××    ×    ×× 
Geogarypidae×   × ××× ×  ×    ×× 
Larcidae      ×  ××        × 
Neobisiidae××× ×××××××××××××××××
Olpiidae×   × ××× ×× ×    ×× 
Syarinidae×   × ×××          × 
Withiidae×   × ××× ×× ××   ×× 
NameUABYMDRU-KGDRU-RUWRU-RUCRU-RUERU-RUNRU-RUSTR-EURTR-ASICYAMAZGEMADZTNLYEGMT
Atemnidae×?         × ××××××× 
Cheiridiidae ? ××× ××    ×  ×    
Cheliferidae×?×××× ××  ××××××××××
Chernetidae×?×××× ××  ××××××××××
Chthoniidae×?×××× ×× ×××××××× ××
Garypidae ?         ××   ×××××
Garypinidae ?         × × ×× ×  
Geogarypidae ?           × ××   ×
Neobisiidae×?××××××××××××××××  ×
Olpiidae ?         ××× ××××××
Syarinidae×? ××× ××      ××   ×
Withiidae ?×××× ××    ××××××××
Legende/Legend
? Checkliste nicht verfügbar No checklist available
× Familienvorkommen bekannt Family documented
(×) Familienvorkommen bekannt, keine Art etabliert Family documented, no species established
  Checkliste enthält Familie nicht Checklist consulted, but family not found
Hinweise zur Nutzung der Tabellen:
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Systematik

Die in Europa heimischen Familien gehören zu den folgenden taxonomischen Kategorien: (Parker 1982) (Harvey 2009)

Quellen

Quellen der Nachweise

  • Carabajal Márquez E, García Carrillo J & Rodríguez Fernández F (2011): Aportaciones al catálogo de los pseudoescorpiones de Andalucía (España) (I) (Arachnida, Pseudoscorpiones). Boletín de la Sociedad Entomológica Aragonesa (S.E.A.) 48, 115–128.
  • Christophoryová J & Jablonski D (2017): New data concerning the distribution of pseudoscorpions in Albania (Pseudoscorpiones: Chernetidae). Natura Croatica 26(1), 117–122, doi:10.20302/nc.2017.26.10.
  • Christophoryová J & Červená M (2020): Apocheiridium ferum (Simon, 1879) (Arachnida, Pseudoscorpiones, Cheiridiidae), a newly recorded genus and species of pseudoscorpion for Hungary. Check List 16 (2), 223–228, doi:10.15560/16.2.223.
  • Christophoryová J, Grula D & Jablonski D (2017): First record of the genus Lamprochernes (Pseudoscorpiones: Chernetidae) in Albania. Biharean Biologist 11(1), 62–64.
  • Christophoryová J, Gruľa D & Krajčovičová K (2017): New records of pseudoscorpions (Arachnida: Pseudoscorpiones) associated with animals and human habitats in Slovakia and the Czech Republic. Arachnologische Mitteilungen/Arachnology Letters 53, S. 67–76, doi:10.5431/aramit5311.
  • Dimitrijević RN & Rađa T (2009): On two new pseudoscorpions (Pseudoscorpiones, Arachnida) from Dalmatia (Croatia). Advances in Arachnology and Developmental Biology 12, 259–266.
  • Dimitrijević RN & Rađa T (2016): On the biodiversity of pseudoscorpions in Croatia: Neobisium curcici (Pseudoscorpiones: Neobisiidae), a new cave-dwelling species from Dalmatia (Croatia). Ecologica Montenegrina 7, 259–265.
  • Dimitrijević RN & Rađa T (2017): Neobisium radjai n. sp. (Neobisiidae: Pseudoscorpiones), a new cave-dwelling pseudoscorpion from Bosnia and Herzegovina. Ecologica Montenegrina 15, 10–16.
  • Fjellberg A & Lissner J (2016): Anthrenochernes stellae Lohmander, 1939 (Pseudoscorpiones , Chernetidae) new to Norway. Norwegian Journal of Entomology 63, S. 4–5.
  • Gardini G, Galli L, Zinni M (2017): Redescription of Geogarypus minor, type species of the genus Geogarypus, and description of a new species from Italy (Pseudoscorpiones: Geogarypidae). The Journal of Arachnology 45 (3), 424–443.
  • Gardini G (2000): Catalogo degli Pseudoscorpioni d’Italia (Arachnida). Fragmenta entomologica 32 (suppl.), 1–181.
  • Gardini G (2009): Chthonius thaleri, a new endogeic species from Venetia, Italy (Pseudoscorpiones: Chthoniidae). Contributions to Natural History 12, 545–550.
  • Gardini G (2013): A revision of the species of the pseudoscorpion subgenus Chthonius (Ephippiochthonius) (Arachnida, Pseudoscorpiones, Chthoniidae) from Italy and neighbouring areas. Zootaxa 3655, S. 1–151.
  • Gardini G (2014): The species of the Chthonius heterodactylus group (Arachnida, Pseudoscorpiones, Chthoniidae) from the eastern Alps and the Carpathians. Zootaxa 3887(2), 101–137, doi:10.11646/zootaxa.3887.2.1.
  • Gardini G (2020): Zaragozachthonius (Pseudoscorpiones, Chthoniidae), a new genus with species in Italy and the Balkan peninsula. Zootaxa 4894 (4), 535–548, doi:10.11646/zootaxa.4894.4.3.
  • Gardini G (2021): The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries. Zootaxa 4987, 1–131, doi:10.11646/zootaxa.4987.1.1.
  • Harvey MS (2011): Pseudoscorpions of the World, version 2.0. Western Australian Museum, Perth.
  • Harvey MS (2013): Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth.
  • Koch L (1873): Übersichtliche Darstellung der europäischen Chernetiden. von Bauer & Raspe Nürnberg, S. 1–68.
  • Kolesnikov VB, Turbanov IS & Gongalsky KB (2019): Two new species false scorpion genus Ephippiochthonius Beier, 1930 (Arachnida: Pseudoscorpiones: Chthoniidae) from the Western Ciscaucasia, Russia. Arthropoda Selecta 28(1), 73–82.
  • Krajčovičová K, Christophoryová J & Mahnert V (2017): Rhacochelifer disjunctus (Pseudoscorpiones: Cheliferidae) new to the fauna of Slovakia. Arachnologische Mitteilungen/Arachnology Letters 53, S. 38–42, doi:10.5431/aramit5306.
  • Krajčovičová K, Ivinskis P, Rimsaite J & Christophoryová J (2020): Chernes similis (Beier, 1932) (Pseudoscorpiones, Chernetidae) new to the fauna of Lithuania. Check List 16(3), 707–710, doi:10.15560/16.3.707.
  • Lissner J (2016): A small study of the Corsican spider and pseudoscorpion fauna (Araneae, Pseudoscorpiones). SPINED 36, S. 5–15.
  • Lissner J (2019): First Records of the Pseudoscorpion Microbisium suecicum Lohmander, 1945 in Denmark with Notes on M. brevifemoratum (Ellingsen, 1903). Newsl. Br. arachnol. Soc. 146, 2–5.
  • Lissner J (2021): External morphology and habitat preferences of Ephippiochthonius tetrachelatus and E. kewi (Pseudoscorpiones: Chthoniidae). Arachnologische Mitteilungen 61, 50–57, doi:10.30963/aramit6108.
  • Mahnert V & Weber D (2013): Pseudoscorpione (Arachnida, Pseudoscorpiones) aus Höhlen des Großherzugtums Luxemburg [in: Weber Dieter (Hrsg) 2013. - Die Höhlenfauna Luxemburgs]. Ferrantia 69, S. 108–114.
  • Muster C & Blick T (2016): Rote Liste und Gesamtartenliste der Pseudoskorpione (Arachnida: Pseudoscorpiones) Deutschlands. Naturschutz und Biologische Vielfalt 70 (4), S. 539–561.
  • Nassirkhani M, Zaragoza JA, Snegovaya N & Chumachenko YA (2020): Description of two new Neobisium (Neobisium) species and redescription of Neobisium (N.) speleophilum from Caucasian Russia, with a key to the Neobisium (Neobisium) species (Arachnida: Pseudoscorpiones) recorded from Russia. Arachnology 18 (5), 449–461.
  • Novák J, Hörweg C (2017): Redescriptions of three species of the genus Neobisium from the Balkans and the Carpathians (Arachnida: Pseudoscorpiones). Acta Zoologica Academiae Scientiarum Hungaricae 63 (3), 277–292, doi:10.17109/AZH.63.3.277.2017.
  • Novák J (2017): Neobisium (N.) tothi sp. nov., a new species from Hungary and Romania, and first records of Neobisium (N.) noricum Beier, 1939 from Hungary (Pseudoscorpiones: Neobisiidae). Turkish Journal of Zoology 41, S. 416–423, doi:10.3906/zoo-1607-19.
  • Pfliegler WP, Schönhofer A, Niedbała W, Vella P, Sciberras A & Vella A (2017): New records of mites (Acari) and harvestmen (Opiliones) from Malta with a preliminary checklist of Maltese Arachnida. Soil Organisms 89 (2), S. 85–110, ISSN 2509-9523.
  • Reboleira AS, Zaragoza JA, Gonçalves F & Oromí P (2013): On hypogean Roncocreagris (Arachnida: Pseudoscorpiones: Neobisiidae) from Portugal, with descriptions of three new species. Zootaxa 3670 (2), 283–299, doi:10.11646/zootaxa.3670.2.11.
  • Reboleira ASPS, Zaragoza JA, Gonçalves F & Oromí P (2012): Lusoblothrus, a new syarinid pseudoscorpion genus (Arachnida) from Portugal, occupying an isolated position within the Holarctic fauna. Zootaxa 3544, S. 52–62, ISSN 1175-5334.
  • Turbanov IS & Kolesnikov VB (2021): Three new hypogean species of the false scorpions genus Ephippiochthonius Beier, 1930 (Arachnida: Pseudoscorpiones: Chthoniidae) from the Crimean Peninsula. Arthropoda Selecta 30(2), 193–204.
  • Turbanov IS, Kolesnikov VB (2020): Two new cave-dwelling species of the false scorpion genus Pseudoblothrus Beier, 1931 (Arachnida: Pseudoscorpiones:Syarinidae) from the Crimean Peninsula. Arthropoda Selecta 29(1), 28–50, doi:10.15298/arthsel.29.1.03.
  • Witkowski ZJ, Król W & Solarz W (eds.) (2003): Carpathian List of Endangered Species. WWF and Institute of Nature Conservation, Polish Academy of Sciences, Vienna-Krakow. ISBN 83-918914-0-2, 84 S.
  • Zaragoza JA & Reboleira AS (2018): Five new hypogean Occidenchthonius (Pseudoscorpiones: Chthoniidae) from Portugal. Journal of Arachnology 46(1), S. 81–103, doi:10.1636/JoA-S-17-031.1.
  • Zaragoza JA & Vadell M (2013): Chthonius (Chthonius) campaneti, a new pseudoscorpion species from Coves de Campanet, Mallorca (Spain). Revista Ibérica de Aracnología nº 23, 33–41.
  • Zaragoza JA (2017): Revision of the Ephippiochthonius complex in the Iberian Peninsula, Balearic Islands and Macaronesia, with proposed changes to the status of the Chthonius subgenera (Pseudoscorpiones, Chthoniidae). Zootaxa 4246 (1), S. 1–221, doi:10.11646/zootaxa.4246.1.1.
  • Ćurčić BPM, Dimitrijević RN & Stojanović DZ (2012): A new epigean pseudoscorpion from east Serbia: Chthonius (Ephippiochthonius) timacensis n. sp. (Chthoniidae, Pseudoscorpiones). Arch. Biol. Sci. 64 (3), 1093–1098, doi:10.2298/ABS1203093C.
  • Ćurčić BPM, Dimitrijević RN & Ćurčić NB (2011): A new cave pseudoscorpion (Psudoscorpiones: Chthoniidae): Chthonius lupinus n. sp. from Bosnia-Herzegovina. Arch. Biol. Sci. 63 (2), 499–505, doi:10.2298/ABS1102499C.
  • Ćurčić BPM, Dimitrijević RN, Makarov SE, Ćurčić SB, Tomić VT, Antić DŽ, Ilić BS & Ćurčić NB (2013): Roncus radgost n. sp., R. jarevid n. sp., and R. crnobog n. sp.: Three new cave dwellers from eastern Serbia (Neobisiidae, Pseudoscorpiones). Arch. Biol. Sci. 65 (2), 751–760, doi:10.2298/ABS1302751Ć.
  • Ćurčić BPM, Dimitrijević RN, Rađa T & Milinčić M (2012): Chthonius (Chthonius) makirina (Chthoniidae, Pseudoscorpiones), a new species from Croatia. Arch. Biol. Sci. 64 (2), 709–714, doi:10.2298/ABS1202709C.
  • Ćurčić BPM, Dimitrijević RN, Rađa T, Makarov SE, Ilić BS (2012): Archaeoroncus, a New Genus of Pseudoscorpions from Croatia (Pseudoscorpiones: Neobisiidae), with Descriptions of Two New Species. Acta Zoologica Bulgarica 64 (4), 333–340.
  • Ćurčić BPM, Dimitrijević RN, Tomić VT, Pecelj M, Ilić BS, Makarov SE, Ćurčić NB & Pešić S (2012): A new epigean false scorpion: Roncus sumadijae n. sp. (Neobisiidae, Pseudoscorpiones) from the Balkan peninsula (western Serbia). Arch. Biol. Sci. 64 (2), 703–708, doi:10.2298/ABS1202703C.
  • Ćurčić BPM, Makarov SE, Rađa T, Ilić BS & Antić DŽ (2012): Roncus meledae n. sp. and Neobisium oculatum n. sp., from the island of Mljet, Dalmatia (Neobisiidae, Pseudoscorpiones). Arch. Biol. Sci. 64 (4), 1567–1576, doi:10.2298/ABS1204567C.
  • Ćurčić BPM, Makarov SE, Ćurčić NB, Ćurčić SB & Tomić VT (2013): Chthonius (Ephippiochthonius) rhizon n. sp.: A new cave false scorpion from the bay of Kotor (Chthoniidae, Pseudoscorpiones). Arch. Biol. Sci. 65 (4), 1547–1551, doi:10.2298/ABS1304547C.
  • Ćurčić BPM, Makarov SE, Ćurčić SB, Antić DŽ & Dimitrijević RN (2012): A new soil pseudoscorpion, Roncus ursi n. sp., from western Serbia (Neobisiidae, Pseudoscorpiones). Arch. Biol. Sci. 64 (1), 379–384, doi:10.2298/ABS1201379C.
  • Ćurčić BPM, Makarov SE, Ćurčić SB, Rađa T, Tomić VT & Ilić BS (2012): A new rare representative of Microchthonius Hadži (Chthoniidae, Pseudoscorpiones) from Dalmatia, Croatia. Acta Zoologica Bulgarica 64 (3), 229–234.
  • Ćurčić BPM, Rađa T & Dimitrijević RN (2012): On two new cave pseudoscorpions, Chthonius (Chthonius) pagus n. sp. (Chthoniidae) and Roncus navalia n. sp. (Neobisiidae), from the island of Pag, Croatia. Arch. Biol. Sci. 64 (4), 1555–1565, doi:10.2298/ABS1204555C.
  • Ćurčić BPM, Rađa T, Dimitrijević R, Vesović N & Ćurčić S (2015): On two new species of Microchthonius Hadži (Pseudoscorpiones, Chthoniidae) from Dalmatia, Croatia. Bul. Shk., Ser. Shk. Nat. 65, 80-93.
  • Ćurčić BPM, Rađa T, Dimitrijević RN & Ćurčić S (2012): On two new cave pseudoscorpions from Dalmatia, Croatia (Chthoniidae and Neobisiidae, Pseudoscorpiones). Arch. Biol. Sci. 64 (3), 1099–1108, doi:10.2298/ABS1203099C.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Makarov SE, Milinčić M & Pecelj JM (2011): Two new pseudoscorpions from the UN administered province of Kosovo and Croatia. Arch. Biol. Sci. 63 (1), 235–244, doi:10.2298/ABS1101235C.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Vesović NR, Ćurčić NB & Ćurčić SB (2021): Roncus sutikvae sp. n. (Pseudoscorpiones: Neobisiidae), a new epigean pseudoscorpion from central Dalmatia (Croatia). Arthropoda Selecta 30(2), 205–215.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Ćurčić NB, Ilić BS & Pecelj JM (2011): Chthonius (Ephippiochthonius) lagadini n. sp. (Chthoniidae, Pseudoscorpiones), a new endemic epigean pseudoscorpion from Macedonia. Arch. Biol. Sci. 63 (4), 1251–1256, doi:10.2298/ABS1104251C.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Ćurčić SB, Makarov SE, Antić DŽ & Ilić BS (2014): A new pseudoscorpion from Bosnia: Roncus bosniensis n. sp. (Neobisiidae, Pseudoscorpiones). Arch. Biol. Sci. 66 (1), 363–368, doi:10.2298/ABS1401363Ć.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Ćurčić SB, Ćurčić NB & Makarov SE (2014): On two new cave species of Pseudoscorpions (Neobisiidae, Pseudoscorpiones) from Herzegovina and Dalmatia. Arch. Biol. Sci. 66 (1), 377–384, doi:10.2298/ABS1401377C.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Ćurčić SB, Ćurčić NB, Makarov SE & Ilić BS (2014): Microchthonius kasteli n. sp. (Chthoniidae, Pseudoscorpiones) — A new cave false scorpion from Croatia (Dalmatia). Arch. Biol. Sci. 66 (1), 437–443, doi:10.2298/ABS1401437C.
  • Ćurčić BPM, Rađa T, Dimitrijević RN, Ćurčić SB, Ćurčić NB and Makarov SE (2013): A new cave pseudoscorpion from Dalmatia — Microchthonius tragurion n. sp. (Chthoniidae, Pseudoscorpiones). Arch. Biol. Sci. 65 (3), 1253–1259, doi:10.2298/ABS1303253C.
  • Ćurčić BPM, Rađa T, Makarov SE, Ćurčić S, Ilić BS & Dimitrijević RN (2011): A cavernicolous pseudoscorpion of the genus Chthonius (Chthonius) C. L. Koch from Dalmatia. Arch. Biol. Sci. 63 (2), 493–497, doi:10.2298/ABS1102493C.
  • Ćurčić BPM, Rađa T, Rađa B, Dimitrijević RN, Tomić VT & Ilić BS (2013): A New Cave Pseudoscorpion from the Adriatic Isle of Šolta (Dalmatia): Microchthonius solentanus n. sp. (Pseudoscorpiones: Chthoniidae). Acta Zoologica Bulgarica 65 (3), 283–287.
  • Ćurčić BPM, Rađa T, Rađa B, Dimitrijević RN, Ćurčić SB, Ćurčić NB & Ilić BS (2014): Chthonius (Globochthonius) daorsoni n. sp. (Chthoniidae, Pseudoscorpiones) — A new cave false scorpion from Bosnia and Herzegovina. Arch. Biol. Sci. 66 (2), 901–906, doi:10.2298/ABS1402901Ć.
  • Ćurčić BPM, Rađa T, Ćurčić SB, Ilić BS, Tomić VT & Makarov SE (2013): Microchthonius elegantissimus n. sp., a new troglobitic pseudoscorpion (Pseudoscorpiones, Chthoniidae) from Croatia. rch. Biol. Sci. 65 (1), 405–410, doi:10.2298/ABS1301405C.
  • Ćurčić BPM, Sarbu SM, Dimitrijević RN & Ćurčić SB (2014): A New Cave Pseudoscorpion from the region of Mangalia (Romania): Chthonius (Ephippiochthonius) borissketi n. sp. (Chthoniidae, Pseudoscorpiones). Arch. Biol. Sci. 66 (2), 955-961, doi:10.2298/ABS1402955C.
  • Ćurčić BPM, Stojanović DZ, Ilić BS & Ćurčić NB (2012): Roncus ivansticae (Neobisiidae, Pseudoscorpiones): A new epigean species from eastern Serbia. Arch. Biol. Sci. 64 (1), 371–377, doi:10.2298/ABS1201371C.
  • Ćurčić BPM, Ćurčić SB, Ćurčić NB, Rađa T & Dimitrijević RN (2011): On two new Pseudoscorpions from Herzegovina. Arch. Biol. Sci. 63 (3), 855–865, doi:10.2298/ABS1103855C.
  • Ćurčić NB & Dimitrijević RN (2016): Checklist of the pseudoscorpions (Arachnida: Pseudoscorpiones) of Montenegro. Ecologica Montenegrina 7, S. 439–450.
  • Ćurčić NB, Dimitrijević RN & Ćurčić S (2020): Checklist of the pseudoscorpions (Arachnida: Pseudoscorpiones) of Serbia. Arthropoda Selecta 29(1), 13–27, doi:10.15298/arthsel.29.1.02.
  • Červená M, Gardini G & Christophoryová J (2021): New contributions to the knowledge of pseudoscorpion diversity (Arachnida, Pseudoscorpiones) of Moldova after more than 30 years. Arachnologische Mitteilungen 61, 45–49, doi:10.30963/aramit6107.
  • Červená M, Gardini G, Jablonski D & Christophoryová J (2021): Checklist of Pseudoscorpions (Arachnida, Pseudoscorpiones) of Albania. Zoological Studies 60:17, 1–25, doi:10.6620/ZS.2021.60-17.
  • Červená M, Krajčovičová K & Christophoryová J (2020): Updated checklist of pseudoscorpions (Arachnida: Pseudoscorpiones) of Central Europe. Arthropoda Selecta 29(2), 219–228.
  • Červená M, Selnekovič D & Christophoryová J (2021): New records of chernetid and cheliferid species (Arachnida: Pseudoscorpiones) from North Macedonia. Natura Croatica 30 (1), 251–256, doi:10.20302/NC.2021.30.16.